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Billy Coen, Former Marine, now is a wanted fugitive によって the military to serve a sentence to death because of a crime, but wasn't his fault. and hes still on the run until the end of the game.

On the way to a prison where they put guys like him to death, a military personel vehicle with prison personel including Billy, was attacked によって evil rotweiler イヌ and some worm like Leeches sent out によって Doctor James Marcus along the Arkay Mountian Railroad.
Billy manages to outrun himself to the nearby Train called the Ecliptic Express, and then meets Rebecca, a S.T.A.R.S. member and a seemingly trust-worthy partner at hand.